In an interview with GameTrailers, Chet Faliszek, the writer for Left 4 Dead confirmed that there will be a demo/beta for both the 360 and PC versions of the game available before release.
Regarding the online play, dedicated servers will be set up so players can experience the least amount of lag as possible. You will still be able to blast zombies solo or with your friends via system link however.
Interview after the break.
-Ryizzel
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